Three linked autosomal loci were studied in squash: flowering time (late vs. early), fruit shape (elongate vs. ovate) and stem length (long vs. short). Pure breeding early flowering, elongated squash with long stems were mated to pure breeding late flowering, ovated squash with short stems. The F1 were testcrossed and the testcross progeny are given below.

Flowering Fruit Stem Number
time shape Length
Late Elongate Short 628
Early Elongate Short 72
Early Elongate Long 1481
Late Elongate Long 448
Late Ovate Long 57
Early Ovate Long 585
Early Ovate Short 332
Late Ovate Short 1399


1. How many progeny did you look at for this experiment?

2. Which locus is in the center?

3. What is the value for R1? Use two decimal places in your answer.

4. What is the value for R2? Use two decimal places in your answer.

5. What is the coefficient of coincidence? Use four decimal places in your answer.